@import url('https://fonts.googleapis.com/css2?family=Manrope:wght@400;500;600;700;800&family=IBM+Plex+Mono:ital,wght@0,400;0,500;1,400&display=swap');
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dt{font-family:var(--mono);font-size:11px;color:var(--dim);margin-bottom:6px}.facts dd{margin:0;font-size:20px}.facts .full{grid-column:1/-1}.facts .full dd{font-size:16px}.source-note{font-size:12px;line-height:1.7;color:var(--dim)}.subspec-list{display:grid;gap:10px}.subspec-row{padding:14px;border:1px solid var(--border);text-align:left;display:block;width:100%}.subspec-row strong{font-family:var(--mono);font-size:13px;font-weight:500;display:block}.subspec-row span{display:block;font-size:13px;color:var(--muted);margin-top:4px}.subspec-row:hover{border-color:var(--gold)}.subspec-note{color:var(--dim);font-size:12px;line-height:1.7}.subspec-detail h3{font-family:var(--mono);font-size:16px;line-height:1.6}.back-button{color:var(--gold);font-size:14px;padding:0 0 14px;text-decoration:underline;text-underline-offset:4px}.sources-panel a{display:block;padding:12px 0;color:var(--gold);text-decoration:underline;overflow-wrap:anywhere;font-size:14px}.sources-panel p{color:var(--muted);font-size:14px;line-height:1.8}.about-content{padding:42px;position:relative;max-width:780px;margin:auto}.about-content h2{font-size:36px;margin-top:10px}.about-content p{color:var(--muted);line-height:1.8}.about-content a:not(.close){display:block;color:var(--gold);text-decoration:underline;margin:14px 0;font-size:14px}.skip{position:fixed;top:-80px;left:10px;z-index:40;padding:12px;background:var(--gold);color:var(--bg)}.skip:focus{top:10px}
@media(min-width:1560px){main{padding-left:48px;padding-right:48px}.grid{gap:28px}.photo{aspect-ratio:1.55}.card-content{padding:22px}.species-title{font-size:27px}.card-stats>div{font-size:15px}.photo-credit{padding-left:22px;padding-right:22px}}
@media(max-width:1180px){.grid{grid-template-columns:repeat(2,minmax(0,1fr))}.layout{grid-template-columns:218px minmax(0,1fr)}.sidebar{padding:28px 16px}main{padding-left:26px;padding-right:26px}.header-note{display:none}.detail-body{padding:30px 26px}.detail-body h2{font-size:32px}}
@media(max-width:780px){.header{height:74px;padding:0 20px}.brand-mark{font-size:25px}.brand-name{font-size:11px}.brand-name small{font-size:9px}.header-actions{gap:12px}.text-button{font-size:12px}.divider{display:none}.theme-button{width:39px}.layout{display:block}.sidebar{display:none;position:fixed;top:74px;left:0;width:280px;height:calc(100vh - 74px);background:var(--panel);z-index:9;box-shadow:var(--shadow)}.sidebar.mobile-open{display:block}.side-foot{margin-top:25px}main{padding:22px 20px 0}.catalog-heading{margin:22px 0}.catalog-heading h1{font-size:42px}.edition{display:none}.search-wrap{padding:0 12px}.search-wrap input{font-size:14px}kbd{display:none}#mobile-taxonomy{display:block;font-size:12px;padding:8px}.filter-line{align-items:flex-start;flex-direction:column;gap:11px;margin:14px 0 20px}.filters{width:100%;flex-wrap:nowrap;overflow-x:auto;padding-bottom:5px}.filters button{white-space:nowrap;flex-shrink:0;font-size:12px;padding:8px 10px}.results{font-size:12px}.grid{gap:20px 14px}.card-content{padding:14px}.species-title{font-size:21px}.card-stats{gap:8px;flex-wrap:wrap}.card-stats>div{font-size:12px}.subspecies-button{font-size:11px}.photo-credit{padding:0 14px 12px;font-size:10px}.card-region{font-size:12px;min-height:36px}.catalog-footer{flex-direction:column;font-size:10px;line-height:1.8}.detail-grid{grid-template-columns:1fr}.detail-visual img{height:auto}.detail-index{display:none}.detail-body{padding:25px}.detail-body h2{font-size:32px}.detail-visual .photo-credit{padding:12px 25px}.close{background:var(--panel);border:1px solid var(--border);top:9px;right:9px}dialog{width:calc(100% - 24px);max-height:calc(100dvh - 24px)}.detail-visual{position:relative}.about-content{padding:30px}.breadcrumbs{font-size:11px}}
@media(max-width:520px){.grid{grid-template-columns:1fr}.photo{aspect-ratio:1.48}.card-content{padding:18px}.species-title{font-size:26px}.card-stats>div{font-size:14px}.card-stats .stat-label{font-size:11px}.card-region{min-height:0;font-size:14px}.latin{font-size:14px}.photo-credit{padding:0 18px 14px;font-size:11px}.genus-label{font-size:11px}.catalog-heading h1{font-size:40px}.header-actions .text-button{display:none}.detail-tabs{gap:19px}.search-wrap input::placeholder{font-size:12px}.detail-body h2{overflow-wrap:anywhere}.facts dd{font-size:19px}.facts{gap:18px 10px}}
@media(prefers-reduced-motion:reduce){*{scroll-behavior:auto!important;transition:none!important}}
.filters button{font-size:.875rem}.filters button span{font-size:.75rem}.results{font-size:.75rem}.search-wrap input{font-size:1rem}.catalog-heading p{font-size:1rem}.latin,.card-region{font-size:.875rem}.card-stats{flex-wrap:wrap;gap:12px}.card-stats>div{font-size:.9375rem}.card-stats .stat-label{font-size:.75rem}.subspecies-button{font-size:.875rem;min-height:28px}.photo-credit{font-size:.75rem}.genus-label{font-size:.75rem}.cat-card{touch-action:pan-y}.subspec-row strong,.subspec-row span{font-size:.875rem}.detail-visual .photo-credit{font-size:.75rem}.sources-panel p{font-size:1rem}.side-foot a{font-size:.875rem}

.cat-day,.map-entry{display:inline-flex;align-items:center;justify-content:center;color:var(--gold);background:transparent;border:1px solid var(--gold);padding:10px 15px;min-height:44px;min-width:152px;white-space:nowrap;font:inherit;font-size:14px;line-height:1.5;text-align:center}.cat-day:hover,.map-entry:hover{background:var(--gold-soft)}.detail-photo{position:relative;touch-action:pan-y}.detail-photo img{display:block}.detail-photo .slide-controls{bottom:18px;left:20px;right:20px}.detail-photo .slide-btn{width:42px;height:42px}.detail-photo-count{color:white;background:#111b;padding:4px 12px;font-family:var(--mono);font-size:12px}.card-region{display:-webkit-box;-webkit-line-clamp:3;-webkit-box-orient:vertical;overflow:hidden;min-height:63px}.card-region svg{display:none}.subspec-detail>.back-button{display:block;margin-bottom:22px}.subspec-detail h3{margin-top:0}.map-options-link{display:inline-block;color:var(--gold);text-decoration:underline;text-underline-offset:4px}.detail-visual img{object-fit:contain;max-width:100%}
@media(max-width:780px){.header{gap:10px}.header-actions{gap:12px}.header-actions .divider{display:none}.detail-photo img{height:auto}.header-note{display:none}}@media(max-width:520px){.cat-day{padding:8px 10px;font-size:12px}.header-actions{gap:8px}.brand-name{font-size:10px}.brand-name small{font-size:8px;letter-spacing:.12em}.brand{gap:10px}.brand-mark{font-size:25px;padding-right:8px}.theme-button{width:38px}.card-region{min-height:0}}

.photo-note{display:block;width:100%;margin-top:6px;color:var(--muted);font-family:var(--font);font-size:12px;line-height:1.5}.photo-credit{flex-wrap:wrap}

@media(max-width:380px){.brand-name{display:none}.header{padding:0 16px}}

.lineage-group{margin-top:18px}.lineage-heading{padding:0 8px 6px;color:var(--muted);font-family:var(--mono);line-height:1.5}.lineage-heading span{display:block;font-size:12px;letter-spacing:.08em}.lineage-heading strong{font-size:14px;font-weight:500;color:var(--text)}

.about-content p,.sources-panel p,.subspec-note{overflow-wrap:anywhere}

.atlas-nav{display:flex;align-items:center;gap:24px;border-bottom:1px solid var(--border);margin-bottom:22px}.atlas-nav a{font-size:14px;padding:10px 0 14px;color:var(--muted)}.atlas-nav a[aria-current]{color:var(--gold);border-bottom:2px solid var(--gold)}.atlas-nav a:hover{color:var(--gold)}.atlas-nav a span{font-size:12px;margin-left:5px}.detail-actions{display:flex;flex-wrap:wrap;gap:12px 24px;margin-top:20px}.detail-actions .back-button{font-size:14px;min-height:38px}

.header-primary-actions{display:flex;align-items:center;gap:12px}
@media(max-width:680px){.header{height:116px;display:grid;grid-template-columns:minmax(0,1fr) auto;grid-template-rows:56px 48px;gap:0 14px;padding:0 18px 12px}.header .brand{grid-row:1;grid-column:1}.header .brand-name{display:block;font-size:11px}.header-actions{display:contents}.header-actions>.text-button,.header-actions>.divider{display:none}.header .theme-button{grid-row:1;grid-column:2;justify-self:end}.header-primary-actions{grid-row:2;grid-column:1/-1;gap:10px}.header-primary-actions .cat-day,.map-entry{flex:1;min-width:0;text-align:center;font-size:13px;padding:10px 12px;min-height:42px}.sidebar{top:116px;height:calc(100dvh - 116px)}}

/* Subspecies reuse the existing atlas card, palette and typography. */
[hidden]{display:none!important}
.parent-info{color:var(--gold);font-size:14px;text-decoration:underline;text-underline-offset:4px;padding:6px 0;margin-left:16px}
.subspecies-card{position:relative;min-width:0;display:flex;flex-direction:column;background:var(--card);border:1px solid var(--border);transition:border-color .2s,transform .2s}
.subspecies-card:hover{border-color:var(--gold);transform:translateY(-3px)}
.subspecies-card .photo{flex:none}
.subspecies-card .card-content{flex:1;display:flex;flex-direction:column}
.subspecies-title{padding:0;text-align:left;font-size:24px;line-height:1.35;font-weight:600;letter-spacing:-.035em;overflow-wrap:anywhere}
.subspecies-title:after{content:'';position:absolute;inset:0;z-index:1}
.subspecies-card .latin{margin:8px 0 16px}
.subspecies-range{margin-top:18px;color:var(--muted)}
.subspecies-range .stat-label{display:block;font-size:11px;color:var(--dim);letter-spacing:.05em}
.subspecies-range p{font-size:14px;line-height:1.8;margin:7px 0 16px}
.subspecies-more{display:block;padding-top:14px;margin-top:auto;border-top:1px solid var(--border);color:var(--gold);font-size:13px}
.subspecies-card .photo-credit{margin-top:8px;position:relative;z-index:2}
.photo-unconfirmed{height:100%;min-height:180px;background:var(--panel);display:flex;flex-direction:column;justify-content:center;align-items:center;gap:12px;padding:28px;text-align:center;color:var(--dim);border-bottom:1px solid var(--border)}
.photo-unconfirmed strong{font-family:var(--font);font-size:18px;line-height:1.45;font-weight:500;color:var(--muted);max-width:230px}
.photo-unconfirmed>span:last-child{font-size:12px;max-width:230px;line-height:1.7}
.subspecies-parent{font-size:13px;color:var(--dim);margin:20px 0 26px}
.subspecies-parent button{display:block;text-align:left;color:var(--gold);padding:5px 0;text-decoration:underline;text-underline-offset:4px;font-size:14px}
.subspecies-detail .detail-body h2{font-size:32px;overflow-wrap:anywhere}
.subspecies-detail .detail-photo .photo-unconfirmed{min-height:340px}
.subspecies-detail .detail-body>.source-note{margin:18px 0 26px}
.subspecies-evidence{border-block:1px solid var(--border);padding:15px 0;color:var(--muted);font-size:13px;line-height:1.8;overflow-wrap:anywhere}
.subspecies-evidence summary{cursor:pointer;color:var(--gold);font-size:14px}
.subspecies-evidence a{display:block;text-decoration:underline;text-underline-offset:4px;color:var(--gold);margin:12px 0}
.subspecies-mini-grid{grid-template-columns:repeat(2,minmax(0,1fr));gap:14px;margin-top:12px}
.subspecies-mini-grid .subspec-row{padding:0}
.subspecies-mini-grid .subspecies-title{font-size:17px}
.subspecies-mini-grid .card-content{padding:14px}
.subspecies-mini-grid .genus-label{display:none}
.subspecies-mini-grid .subspecies-range .stat-label{font-size:10px}
.subspecies-mini-grid .subspecies-range p{font-size:12px;line-height:1.6;display:-webkit-box;-webkit-line-clamp:4;-webkit-box-orient:vertical;overflow:hidden}
.subspecies-mini-grid .photo-credit{font-size:10px;padding:0 14px 12px}
.subspecies-mini-grid .photo-unconfirmed{min-height:130px;padding:16px;gap:8px}
.subspecies-mini-grid .photo-unconfirmed strong{font-size:14px}
.subspecies-mini-grid .photo-unconfirmed>span{font-size:10px}
.subspecies-mini-grid .subspecies-more{font-size:11px}
@media(max-width:780px){.parent-info{display:block;margin-left:0}.subspecies-detail .detail-photo .photo-unconfirmed{min-height:220px}.subspecies-detail .detail-body h2{font-size:28px}}
@media(max-width:520px){.subspecies-mini-grid{grid-template-columns:1fr}.subspecies-mini-grid .photo{aspect-ratio:1.6}.subspecies-mini-grid .subspecies-title{font-size:22px}.subspecies-mini-grid .subspecies-range p{font-size:14px}.subspecies-card .genus-label{line-height:1.6}.subspecies-title{font-size:25px}}
/* Preserve complete licensed frames and photographer signatures. */
.photo>img[data-preserve-frame]{object-fit:contain!important;filter:none!important;transform:none!important}

.subspecies-description{font-size:14px;line-height:1.8;color:var(--muted);margin:14px 0 22px}.subspecies-mini-grid .subspecies-description{font-size:12px;line-height:1.65;margin:8px 0 15px}.subspecies-parent{margin:18px 0 20px}.conservation-note{font-size:13px;line-height:1.7;color:var(--muted)}.detail-badge[data-status=unknown]{white-space:normal;line-height:1.6}.sources-panel{padding:8px 0 20px}.sources-panel>a{display:inline-block;line-height:1.7}.subspecies-mini-grid .badge{font-size:10px;padding:4px 8px}.subspecies-card .badge{z-index:2;pointer-events:none}

.about-content>.close{display:grid;place-items:center;text-decoration:none;margin:0}

.catalog-footer{gap:12px;flex-wrap:wrap}